Composition of species richness (a) and abundance (b) in four functional guilds. Richness composition is not significantly different between years (χ
2 test, df = 3, P > 0.05), but abundance composition is significantly different between years (P < 0.05). Different letters above the bars indicate significant difference between years. Guilds were defined as follows: FGF, forest ground forager; SLD, soil and litter dweller; OF, open-land forager; FVF, forest vegetation forager. Definition for the guilds is shown in the text.